Wille, Alvaro. “Coleoptera: Scarabaeidae Observations on the Behavior of a Tropical Rain Forest Dung Beetle, <i>Megathoposoma candezei</i≫ (Harold)”. Revista de Biología Tropical 21, no. 1 (July 1, 1973): 41–57. Accessed June 26, 2025. https://archivo.revistas.ucr.ac.cr/index.php/rbt/article/view/27322.